The Grizzly bear (Ursus arctos horribilis) was first described as grisley by Meriwether Lewis and William Clark – the name could mean "grizzly" (i.e., "grizzled"—that is, with grey-tipped hair) or "grisly" ("fear-inspiring", now usually "gruesome"). It is one of the largest subspecies of Brown Bear, only being beaten by the Kamchatka brown bears and the Kodiak bears. Grizzly bears are powerful, top-of-the-food-chain predators, yet much of their diet consists of nuts, berries, fruit, leaves, and roots. Bears also eat other animals, from rodents to moose.